What is a Cetacean?
Lesson 1 · Mammals that chose the ocean
Articles and studies
- A diminutive new basilosaurid whale reveals the trajectory of the cetacean life histories during the Eocene (2023) – Communications Biology
- Basilosaurus, the “King of Lizards” (2020) – Whale Scientists
- Hippos and Whales: Unlikely Cousins (2017) – Royal Ontario Museum
- Science Word of the Day: Involucrum (2015) – National Geographic
- Pakicetus (2022) – EBSCO Research Starters
- From Land to Water: the Origin of Whales, Dolphins, and Porpoises (2009) – Evolution: Education and Outreach
- New Protocetid Whale from the Middle Eocene of Pakistan: Birth on Land, Precocial Development, and Sexual Dimorphism (2009) – PLoS ONE
- Whales originated from aquatic artiodactyls in the Eocene epoch of India (2007) – Nature
- The Origin of Whales and the Power of Independent Evidence (2000) – NCSE
- Phylogenetic relationships among cetartiodactyls based on insertions of short and long interspersed elements: Hippopotamuses are the closest extant relatives of whales (1999) – PNAS
- The evolution of whales – Understanding Evolution, UC Berkeley
Books
- Encyclopedia of Marine Mammals, 3rd edition (2018) – Würsig, Thewissen & Kovacs (eds.)
- Spying on Whales (2018) – Nick Pyenson
- The Cultural Lives of Whales and Dolphins (2015) – Hal Whitehead & Luke Rendell
Lesson 2 · Built for water
Articles and studies
- A Whale of a Tail: Unraveling the evolutionary secrets of how whales and dolphins adapted their backbones for aquatic life (2024) – Harvard University
- The Biological Productivity of the Ocean (2012) – Nature Education
- Species review of Cuvier’s beaked whale, Ziphius cavirostris (2011) – International Whaling Commission
- 5.1 Properties of Water – Introduction to Oceanography
- Properties of Water – ResearchGate
- Whales – Smithsonian Ocean
- The loudest voice in the animal kingdom – BBC Earth
- Do whales and dolphins have hair? – Whale and Dolphin Conservation
- Mammal – ScienceDirect Topics
Books
- Encyclopedia of Marine Mammals, 3rd edition (2018) – Würsig, Thewissen & Kovacs (eds.)
- The Cultural Lives of Whales and Dolphins (2015) – Hal Whitehead & Luke Rendell
- Cetacean Societies: Field Studies of Dolphins and Whales (2000) – Mann, Connor, Tyack & Whitehead (eds.)
Lesson 3 · Two great lineages
Articles and studies
- Cetaceans Have Complex Brains for Complex Cognition (2007) – PLoS Biology
- Intro to Odontocetes & Mysticetes (2017) – Ocean Conservation Research
- Vaquita – NOAA Fisheries
- Intelligence, Cognition and Emotion in Cetaceans – The Whale Sanctuary Project
- Are Whales Sentient? The Scientific Evidence (2026) – Biology Insights
Books
- Encyclopedia of Marine Mammals, 3rd edition (2018) – Würsig, Thewissen & Kovacs (eds.)
- Cetacean Societies: Field Studies of Dolphins and Whales (2000) – Mann, Connor, Tyack & Whitehead (eds.)
Lesson 4 · Underwater living
Articles and studies
- Nursing Behavior in Sperm Whales (Physeter macrocephalus) (2023) – Animal Behavior and Cognition
- Fight or flight: antipredator strategies of baleen whales (2008) – Mammal Review
- Cetacean sleep: An unusual form of mammalian sleep (2008) – Neuroscience and Biobehavioral Reviews
- Lactation in Whales and Dolphins: Evidence of Divergence Between Baleen- and Toothed-Species (1997) – Journal of Mammary Gland Biology and Neoplasia
- [TITLE MISSING] – PubMed Central
- Blue Whale Life Cycle: Birth, Growth, and Lifespan (2026) – Science Insights
- Intelligence, Cognition and Emotion in Cetaceans – The Whale Sanctuary Project
Books
- Encyclopedia of Marine Mammals, 3rd edition (2018) – Würsig, Thewissen & Kovacs (eds.)
The Great Filter Feeders
Lesson 1 · The blue whale
Articles and studies
- Colossal ears? How baleen whales hear low-frequency sound (2025) – Hearing Research
- Blue whales (2020) – Current Biology
- Insight into the kinematics of blue whale surface foraging through drone observations and prey data (2020) – PeerJ
- Balaenoptera musculus (Blue Whale) (2018) – IUCN Red List
- A pulsed-air model of blue whale B call vocalizations (2017) – Scientific Reports
- Worldwide decline in tonal frequencies of blue whale songs (2009) – Endangered Species Research
- Studying How Cetaceans Use Sound to Explore Their Environment (1997) – Perspectives in Ethology
- Blue Whale – NOAA Fisheries
- Blue whale – International Whaling Commission
Lesson 2 · The humpback whale
Articles and studies
- Southern Ocean humpback whales are shifting to an earlier return migration (2025) – Scientific Reports
- Study finds humpback whales only ones capable of bubble-net feeding (2025) – University of Hawaiʻi
- Whales always had voices. Roger Payne helped the world hear them (2023) – Greenpeace
- Why Send Whale Song Into Space? (2017) – Literary Hub
- Humpback whales interfering when mammal-eating killer whales attack other species: Mobbing behavior and interspecific altruism? (2017) – Marine Mammal Science
- Cultural revolution in whale songs (2000) – Nature
- How do the ventral grooves of rorquals function? – Baleines en direct
Books
- Encyclopedia of Marine Mammals, 3rd edition (2018) – Würsig, Thewissen & Kovacs (eds.)
- The Cultural Lives of Whales and Dolphins (2015) – Hal Whitehead & Luke Rendell
Lesson 3 · Right whales
Articles and studies
- Extreme longevity may be the rule not the exception in Balaenid whales (2024) – Science Advances
- Decadal decline in maternal body condition of a Southern Ocean capital breeder (2023) – Scientific Reports
- Saving the North Atlantic right whale in a changing ocean: Gauging scientific and law and policy responses (2020) – Ocean & Coastal Management
- Remote Acoustic Monitoring of North Atlantic Right Whales (Eubalaena glacialis) Reveals Seasonal and Diel Variations in Acoustic Behavior (2014) – PLoS ONE
- Swim Speed, Behavior, and Movement of North Atlantic Right Whales (Eubalaena glacialis) in Coastal Waters of Northeastern Florida, USA (2013) – PLoS ONE
- Dangerous dining: surface foraging of North Atlantic right whales increases risk of vessel collisions (2011) – Biology Letters
- Observations of a Female North Atlantic Right Whale in Simultaneous Copulation with Two Males: Supporting Evidence for Sperm Competition (2005) – Aquatic Mammals
- 10 Things You Should Know About North Atlantic Right Whales – NOAA Fisheries
- Protecting North Atlantic right whales from collisions with vessels in the Gulf of St. Lawrence – Transport Canada
Books
- Encyclopedia of Marine Mammals, 3rd edition (2018) – Würsig, Thewissen & Kovacs (eds.)
Lesson 4 · Gray whales
Articles and studies
- An extremely rare gray whale was spotted in the Atlantic. Scientists say it’s only the beginning. (2024) – National Geographic
- Fish Feeding and Rapid Foraging Behavior Switching by Gray Whales (Eschrichtius robustus) in California (2024) – Aquatic Mammals
- Effects of vessel traffic and ocean noise on gray whale stress hormones (2022) – Scientific Reports
- Natal origin of Namibian grey whale implies new distance record for in-water migration (2021) – Biology Letters
- Migrating eastern North Pacific gray whale call and blow rates estimated from acoustic recordings, infrared camera video, and visual sightings (2019) – PLoS ONE
- Tongue and Hyoid Musculature and Functional Morphology of a Neonate Gray Whale (Cetacea, Mysticeti, Eschrichtius robustus) (2015) – The Anatomical Record
- What Happened to Gray Whales during the Pleistocene? The Ecological Impact of Sea-Level Change on Benthic Feeding Areas in the North Pacific Ocean (2011) – PLoS ONE
- Gray whales may increase feeding opportunities for avian benthivores (2008) – Marine Ecology Progress Series
- Fight or flight: antipredator strategies of baleen whales (2008) – Mammal Review
- Side-scan sonar estimates of the utilization of gray whale feeding grounds along Vancouver Island, Canada (1986) – R. Kvitek & J. Oliver
Lesson 5 · The bowhead whale
Articles and studies
- DNA repair and anti-cancer mechanisms in the long-lived bowhead whale (2023) – bioRxiv (preprint)
- Accuracy of the Aspartic Acid Racemization Technique in Age Estimation of Mammals and the Influence of Body Temperature (2018) – NAMMCO Scientific Publications
- Oral cavity hydrodynamics and drag production in Balaenid whale suspension feeding (2017) – Semantic Scholar
- Insights into the Evolution of Longevity from the Bowhead Whale Genome (2015) – Cell Reports
- The transcriptome of the bowhead whale Balaena mysticetus reveals adaptations of the longest-lived mammal (2014) – Aging
- Hydrodynamic and Sensory Factors Governing Response of Copepods to Simulated Predation by Balaenid Whales (2012) – Semantic Scholar
- Models of hydrodynamic flow in the bowhead whale filter feeding apparatus (2004) – Journal of Experimental Biology
- Observations on the Ice-Breaking and Ice Navigation Behavior of Migrating Bowhead Whales (Balaena mysticetus) near Point Barrow, Alaska, Spring 1985 (1988) – JSTOR
Lesson 6 · Filter feeding
Articles and studies
- How Baleen Whales Feed: The Biomechanics of Engulfment and Filtration (2017) – Annual Review of Marine Science
- Metabolic Expenditures of Lunge Feeding Rorquals Across Scale: Implications for the Evolution of Filter Feeding and the Limits to Maximum Body Size (2012) – PLoS ONE
- Mechanics, hydrodynamics and energetics of blue whale lunge feeding: efficiency dependence on krill density (2011) – Journal of Experimental Biology
Toothed Whales
Lesson 1 · Dolphins
Articles and studies
- Some Dolphins Use Tools to Hunt, Revealing Insights Into Cultural Transmission (2025) – Georgetown University
- Episodic-like memory in common bottlenose dolphins (2022) – Current Biology
- Evidence that bottlenose dolphins can communicate with vocal signals to solve a cooperative task (2021) – Royal Society Open Science
- Fine-scale sociality reveals female–male affiliations and absence of male alliances in bottlenose dolphins (Tursiops truncatus) in the Shannon Estuary, Ireland (2019) – Marine Mammal Science
- Decades-long social memory in bottlenose dolphins (2013) – Proceedings of the Royal Society B
- Bottlenose dolphins can use learned vocal labels to address each other (2013) – PNAS
- Cognitive skills in bottlenose dolphin communication (2013) – Trends in Cognitive Sciences
- Dolphin cognition (2004) – Current Biology
Books
- The Animal Ethics Reader (2003) – Armstrong & Botzler (eds.)
Lesson 2 · Sperm whale
Articles and studies
- Sperm whale clans and human societies (2024) – Royal Society Open Science
- Contextual and combinatorial structure in sperm whale vocalisations (2024) – Nature Communications
- Sperm Whale Phonetic Alphabet Proposed for the First Time (2024) – Project CETI
- Deep Machine Learning Techniques for the Detection and Classification of Sperm Whale Bioacoustics (2019) – Scientific Reports
- Diving and Feeding Behaviour of Sperm Whales (Physeter macrocephalus) in the Northwestern Mediterranean Sea (2004) – Aquatic Mammals
- The face that sank the Essex: potential function of the spermaceti organ in aggression (2002) – Journal of Experimental Biology
- Research – Project CETI
Lesson 3 · Beaked whales
Articles and studies
- Navigating the unknown: assessing anthropogenic threats to beaked whales, family Ziphiidae (2024) – Royal Society Open Science
- Behavioural Ecology Traits of Elusive Deep-Diver Whales Unravel a Complex Social Structure Influenced by Female Philopatry and Defence Polygyny (2022) – Frontiers in Marine Science
- Fear of Killer Whales Drives Extreme Synchrony in Deep Diving Beaked Whales (2020) – Scientific Reports
- Diving behaviour of Cuvier’s beaked whales (Ziphius cavirostris) off Cape Hatteras, North Carolina (2019) – Royal Society Open Science
- Advances in research on the impacts of anti-submarine sonar on beaked whales (2019) – Proceedings of the Royal Society B
- Microplastic and macroplastic ingestion by a deep diving, oceanic cetacean: The True’s beaked whale Mesoplodon mirus (2015) – Environmental Pollution
- No mass strandings since sonar ban (2013) – Nature
- Beaked whales respond to simulated and actual navy sonar (2011) – PLoS ONE
- Correlating military sonar use with beaked whale mass strandings: what do the historical data show? (2009) – Aquatic Mammals
- Extreme diving of beaked whales (2006) – Journal of Experimental Biology
- Elements of beaked whale anatomy and diving physiology and some hypothetical causes of sonar-related stranding (2006) – Journal of Cetacean Research and Management
- Understanding the impacts of anthropogenic sound on beaked whales (2005) – Journal of Cetacean Research and Management
Lesson 4 · Pilot whales
Articles and studies
- Life History Parameters, Sociobiology, and Reproductive Strategies of Pilot Whales (2023) – Sex in Cetaceans (book chapter)
- Ritual Slaughter: The Tradition of Pilot Whale Hunting on the Faroe Islands (2021) – Frontiers in Veterinary Science
- Underlying environmental factors and geomagnetic fields linked to 45 pilot whales Globicephala macrorhynchus stranding in Modung white beach, Indonesian coast on 19th February 2021 (2021) – bioRxiv (preprint)
- Vocal repertoire of long-finned pilot whales (Globicephala melas) in northern Norway (2017) – Journal of the Acoustical Society of America
- The Mass Stranding Event of Long-finned Pilot Whales at Golden Bay in February 2017 (2017) – Kokako
- Mobbing-like behavior by pilot whales towards killer whales: a response to resource competition or perceived predation risk? (2015) – Acta Ethologica
- Pilot Whales Attracted to Killer Whale Sounds: Acoustically-Mediated Interspecific Interactions in Cetaceans (2012) – PLoS ONE
- Satellite tracking of long-finned pilot whales (Globicephala melas) following stranding and release in Tasmania, Australia (2012) – Wildlife Research
- The role of synchronized swimming as affiliative and anti-predatory behavior in long-finned pilot whales (2012) – Behavioural Processes
- Synchronous breathing by pilot whales (2012) – Marine Mammal Science
Lessons 5–6 · Porpoises & River dolphins
Articles and studies
- Estimating the abundance of the critically endangered Baltic Proper harbour porpoise (Phocoena phocoena) population using passive acoustic monitoring (2022) – Ecology and Evolution
- A case of playful interaction between Bolivian river dolphins with a Beni anaconda (2022) – JSTOR
- River Dolphins Spotted in Rare Playful Interaction With a Beni Anaconda (2022) – Smithsonian Magazine
- Polluted porpoises: Generational transfer of organic contaminants in harbour porpoises from the southern North Sea (2021) – Science of the Total Environment
- Interacting effects of vessel noise and shallow river depth elevate metabolic stress in Ganges river dolphins (2019) – Scientific Reports
- Porpoises the World Over: Diversity in Behavior and Ecology (2019) – Ethology and Behavioral Ecology of Odontocetes (book chapter)
- High field metabolic rates of wild harbour porpoises (2018) – Journal of Experimental Biology
- Morphological convergence in ‘river dolphin’ skulls (2017) – PeerJ
- Ultra-High Foraging Rates of Harbor Porpoises Make Them Vulnerable to Anthropogenic Disturbance (2016) – Current Biology
- The Neocortex of Indian River Dolphins (Genus Platanista): Comparative, Qualitative and Quantitative Analysis (2016) – Brain, Behavior and Evolution
- Do Porpoises Choose Their Associates? A New Method for Analyzing Social Relationships among Cetaceans (2011) – PLoS ONE
- Coexistence of Fisheries with River Dolphin Conservation (2010) – Conservation Biology
- Bye Baiji? (2007) – Current Biology
- River Dolphins Initiative – WWF
- Myths and Legends of the Amazon’s Pink River Dolphin – South America Vacations
Lesson 7 · Echolocation
Articles and studies
- For Dolphins, Echolocation May Be More Like ‘Touching’ Than ‘Seeing’ (2025) – Scientific American
- Adaptive echolocation behavior of bats and toothed whales in dynamic soundscapes (2023) – Journal of Experimental Biology
- Echolocating toothed whales use ultra-fast echo-kinetic responses to track evasive prey (2021) – eLife
- A bigger nose, a bigger bang: size matters for echolocating toothed whales (2018) – Aarhus University
- Clicking in Shallow Rivers: Short-Range Echolocation of Irrawaddy and Ganges River Dolphins in a Shallow, Acoustically Complex Habitat (2013) – PLoS ONE
- The Acoustic World of Harbor Porpoises – American Scientist
- Possible limitations of dolphin echolocation: a simulation study based on a cross-modal matching experiment – no link
Lesson 8 · Orca
Articles and studies
- Why these orcas are wearing salmon as hats (again) (2024) – National Geographic
- Ancestry testing of “Old Tom,” a killer whale central to mutualistic interactions with human whalers (2023) – Journal of Heredity
- These orcas control the waves to hunt. It’s spine-tingling to watch. (2023) – National Geographic
- Independent acoustic variation of the higher- and lower-frequency components of biphonic calls can facilitate call recognition and social affiliation in killer whales (2020) – PLoS ONE
- Genome-culture coevolution promotes rapid divergence of killer whale ecotypes (2016) – Nature Communications
- The Cooperation of Humans and Killer Whales (Orcinus orca): The Application of a Simple Fuzzy Rule-Based Model to a Historical System (2015) – Academia.edu
- Killer whale ecotypes: Is there a global model? (2013) – Biological Reviews
- Cultural traditions and the evolution of reproductive isolation: ecological speciation in killer whales? (2012) – Biological Journal of the Linnean Society
- Antarctic Peninsula killer whales (Orcinus orca) hunt seals and a penguin on floating ice (2012) – Marine Mammal Science
- Cross-cultural and cross-ecotype production of a killer whale ‘excitement’ call suggests universality (2011) – Naturwissenschaften
- Killer whales (Orcinus orca) feeding on schooling herring (Clupea harengus) using underwater tail-slaps: kinematic analyses of field observations (2000) – Journal of Experimental Biology
- Development of intentional stranding hunting techniques in killer whale (Orcinus orca) calves at Crozet Archipelago (1995) – Canadian Journal of Zoology
- Surface and underwater observations of cooperatively feeding killer whales in northern Norway (1993) – Canadian Journal of Zoology
- Intentional stranding apprenticeship and social play in killer whales (Orcinus orca) (1991) – Canadian Journal of Zoology
- Acoustic behaviour of resident killer whales (Orcinus orca) off Vancouver Island, British Columbia (1989) – Canadian Journal of Zoology
Books
- Endangered Orcas: The Story of the Southern Residents (2019) – Monika Wieland Shields
Lives in the Ocean
Lesson 1 · Migration
Articles and studies
- How Marine Mammals Navigate – A Review (2026) – University of British Columbia, Working Paper
- Migrating baleen whales transport high-latitude nutrients to tropical and subtropical ecosystems (2025) – Nature Communications
- Whales Don’t Ask for Directions: Here’s How They Always Know the Way (2025) – Whale Scientists
- Monitoring whale migration from space (2025) – WWF Arctic
- Whisperers of whales wander: A directional statistical investigation of whales’ migration influenced by geomagnetic, ocean current, and celestial cues (2025) – Journal for Nature Conservation
- Timing is everything: Drivers of interannual variability in blue whale migration (2020) – Scientific Reports
- Gray whales strand more often on days with increased levels of atmospheric radio-frequency noise (2020) – Current Biology
- Straight as an arrow: humpback whales swim constant course tracks during long-distance migration (2011) – Biology Letters
- Evidence from strandings for geomagnetic sensitivity in cetaceans (1986) – Journal of Experimental Biology
- Gray Whale Migration – NOAA Ocean Today
Books
- Encyclopedia of Marine Mammals, 3rd edition (2018) – Würsig, Thewissen & Kovacs (eds.)
Lesson 2 · Ecosystem engineers
Articles and studies
- Migrating baleen whales transport high-latitude nutrients to tropical and subtropical ecosystems (2025) – Nature Communications
- The Great Whale Conveyor Belt: Earth’s Largest Mammals Keep Oceans Thriving (2025) – Whale Scientists
- Opportunities, research gaps, and risks in allogenic ecosystem engineer mimicry (2025) – PubMed Central
- Whales and Carbon Sequestration: Can Whales Store Carbon? (2024) – NOAA Fisheries
- Whales are the engineers of their ecosystems (2021) – Whale Scientists
- Whales are Vital for Marine Ecosystem Health (2016) – International Whaling Commission
- Whales as marine ecosystem engineers (2014) – Frontiers in Ecology and the Environment
- Ecosystem Engineers – ScienceDirect Topics
- Blue Carbon – NOAA Office for Coastal Management
Lesson 3 · Diving
Articles and studies
- How To Study The Deepest Diving Whales? Use Chemistry! (2024) – Whale Scientists
- How did whales become the world’s deepest-diving mammals? (2021) – Whale Scientists
- Extreme diving in mammals: first estimates of behavioural aerobic dive limits in Cuvier’s beaked whales (2020) – Journal of Experimental Biology
- The advantages of diving deep: Fin whales quadruple their energy intake when targeting deep krill patches (2019) – Functional Ecology
- The CROW: Campus Research and Observational Writings, Volume Two (2017) – University of Washington Bothell
- Adaptations 6: Problems with pressure (2012) – The University of Western Australia
- Deadly diving? Physiological and behavioural management of decompression stress in diving mammals (2012) – Proceedings of the Royal Society B
Books
- Encyclopedia of Marine Mammals, 3rd edition (2018) – Würsig, Thewissen & Kovacs (eds.)
Lesson 4 · Culture
Articles and studies
- Animal culture: conservation in a changing world (2025) – Philosophical Transactions of the Royal Society B
- Culture and conservation in baleen whales (2025) – Philosophical Transactions of the Royal Society B
- Cultural Transmission, Evolution, and Revolution in Vocal Displays: Insights From Bird and Whale Song (2020) – Frontiers in Psychology
- Dolphin girl gang uses sponges to catch prey and create a new culture (2020) – Whale Scientists
- The interplay between social networks and culture: theoretically and among whales and dolphins (2013) – Philosophical Transactions of the Royal Society B
- Spontaneous human speech mimicry by a cetacean (2012) – Current Biology
Books
- The Cultural Lives of Whales and Dolphins (2015) – Hal Whitehead & Luke Rendell
Under Threat
Lesson 1 · Whaling
Articles and studies
- Historical Review of Whaling (2020) – Springer (book chapter)
- Too Much Is Never Enough: The Cautionary Tale of Soviet Illegal Whaling (2014) – Marine Fisheries Review
- A Review of Changes in the Use of Whale Resources over Time in Japan, with a Specific Example of the Hand-harpoon Fishery of Nago, Okinawa Prefecture (2013) – Semantic Scholar
- A taxonomy of world whaling: operations, eras, and data sources (2003) – NOAA
- Modern Eskimo whaling in the Alaskan Arctic (1988) – IEEE Xplore
- The Technology of Whaling in Australian Waters in the 19th Century (1983) – Semantic Scholar
- The Development of Indigenous Whaling: Prehistoric and Historic Contexts – ResearchGate
- Commercial whaling – International Whaling Commission
Books
- Encyclopedia of Marine Mammals, 3rd edition (2018) – Würsig, Thewissen & Kovacs (eds.)
Lesson 2 · Bycatch and entanglement
Articles and studies
- The State of World Fisheries and Aquaculture 2024: Fisheries and aquaculture employment (2024) – FAO
- How ‘dolphin safe’ is canned tuna, really? (2021) – National Geographic
- Bycatch in gillnet fisheries threatens Critically Endangered small cetaceans and other aquatic megafauna (2019) – Endangered Species Research
- Entanglement is a costly life-history stage in large whales (2017) – Ecology and Evolution
- Welfare Implications of Cetacean Bycatch and Entanglements (2017) – Marine Mammal Welfare (book chapter)
- The Painful Side of Trap and Fixed Net Fisheries: Chronic Entanglement of Large Whales (2012) – Journal of Marine Biology
- Lethal entanglement in baleen whales (2011) – Diseases of Aquatic Organisms
- Bycatch – International Whaling Commission
Lesson 3 · Noise pollution
Articles and studies
- Pilot whales shout louder to be heard over noise pollution (2026) – Journal of Experimental Biology
- Avoidance, confusion or solitude? Modelling how noise pollution affects whale migration (2024) – Movement Ecology
- Approaches to reduce noise from ships operating in important killer whale habitats (2019) – Marine Pollution Bulletin
- Evolution of Marine Noise Pollution Management (2015) – Aquatic Mammals
- Evidence that ship noise increases stress in right whales (2012) – Proceedings of the Royal Society B
- The impact of ocean noise pollution on marine biodiversity (2008) – Lindy Weilgart
Lesson 4 · Plastic and chemical pollution
Articles and studies
- Plastics – the fast Facts 2024 (2024) – Plastics Europe
- Chemical Pollution and Diseases of Marine Mammals (2023) – Marine Mammals (book chapter)
- 7 Solutions To Ocean Plastic Pollution (2023) – Oceanic Society
- Predicting global killer whale population collapse from PCB pollution (2018) – Science
- Evidence that the Great Pacific Garbage Patch is rapidly accumulating plastic (2018) – Scientific Reports
- Population growth is limited by nutritional impacts on pregnancy success in endangered Southern Resident killer whales (Orcinus orca) (2017) – PLoS ONE
- Exposure to a mixture of toxic chemicals: implications for the health of endangered southern resident killer whales (2016) – NOAA
- The impact of debris on marine life (2015) – Marine Pollution Bulletin
- As main meal for sperm whales: Plastics debris (2013) – Marine Pollution Bulletin
- Cetaceans and Marine Debris: The Great Unknown (2012) – Journal of Marine Biology
- Editorial: Plastic Pollution – An Ocean Emergency – ResearchGate
- Global plastics production – Our World in Data
- Facts – Seaspiracy
Lesson 5 · Climate change
Articles and studies
- Potential synergistic impacts of climate change and anthropogenic activity on Arctic marine mammals (2026) – Environmental Reviews
- The Ocean and Climate Change (2025) – NASA Science
- Climate change and cetacean health: impacts and future directions (2022) – PubMed
- Climate change-accelerated ocean biodiversity loss & associated planetary health impacts (2022) – ScienceDirect
- Rapid climate-driven circulation changes threaten conservation of endangered North Atlantic right whales (2019) – Oceanography
- Living Planet Index (2016) – WWF
- ‘The Blob’ invades Pacific, flummoxing climate experts (2015) – Science
- Living Blue Planet Report (2015) – WWF
- Arctic amplification dominated by temperature feedbacks in contemporary climate models (2014) – Nature Geoscience
- Processes and impacts of Arctic amplification: A research synthesis (2011) – Global and Planetary Change
- Loss of Arctic sea ice causing punctuated change in sightings of killer whales (Orcinus orca) over the past century (2009) – Ecological Applications
- Ocean acidification: the other CO2 problem (2009) – Annual Review of Marine Science
- Potential effects of climate change on marine mammals (2006) – Oceanography and Marine Biology: An Annual Review
- Long-term decline in krill stock and increase in salps within the Southern Ocean (2004) – Nature
- How climate change impacts the ocean – IFAW
- How Narwhals Find Food: Hunting Beneath Arctic Sea Ice – Narwhal.org
Conservation in Action
Lesson 1 · The humpback recovery
Articles and studies
- Changes in age-related sexual selection in a humpback whale population recovering from exploitation (2026) – Current Biology
- From Harpoons to Hope – The Return of Humpback Whales (2025) – Greenpeace Australia Pacific
- Why humpback whales are coming back (2025) – EarthSky
- Bellwethers of change: population modelling of North Pacific humpback whales from 2002 through 2021 reveals shift from recovery to climate response (2024) – Royal Society Open Science
- Humpback Whales Have Made a Remarkable Recovery, Giving Us Hope for the Planet (2020) – TIME
- Humpback whales make stunning comeback in southern Africa (2019) – National Geographic
- Post-whaling Recovery: The Beautiful Story of Humpbacks (2019) – Baleines en direct
- Assessing the recovery of an Antarctic predator from historical exploitation (2019) – Royal Society Open Science
- High pregnancy rates in humpback whales (Megaptera novaeangliae) around the Western Antarctic Peninsula, evidence of a rapidly growing population (2018) – Royal Society Open Science
- Humpback Whales: An Endangered Species Act Success Story? (2016) – NOAA Fisheries
- The whale that inspired Greenpeace (2013) – OUPblog – Frank Zelko
- Humpback whale – International Whaling Commission
- Humpback whale – Center for Biological Diversity
- The recovery of North Atlantic right whales, Eubalaena glacialis, has been constrained by human-caused mortality (2018) – Royal Society Open Science
Lesson 2 · The vaquita
Articles and studies
- Saving the Vaquita (2025) – Sea Shepherd
- Results of the 2025 Vaquita Monitoring Effort (2025) – Sea Shepherd
- Chapter 13 – Vaquita Phocoena sinus Norris and McFarland, 1958 (2025) – ScienceDirect (book chapter)
- Addressing Illegal Transnational Trade of Totoaba and Its Role in the Possible Extinction of the Vaquita (2023) – Journal of International Wildlife Law & Policy
- A field effort to capture critically endangered vaquitas Phocoena sinus for protection from entanglement in illegal gillnets (2019) – Endangered Species Research
- Conservation of the vaquita Phocoena sinus (2006) – Mammal Review
- What is killing the vaquita? – Porpoise Conservation Society
- Vaquita – NOAA Fisheries
Books
- Encyclopedia of Marine Mammals, 3rd edition (2018) – Würsig, Thewissen & Kovacs (eds.)
Lesson 3 · Marine protected areas
Articles and studies
- A New Era for Ocean Protection (2026) – The Nature Conservancy
- What is the High Seas Treaty and Why is it Important? (2026) – United Nations University
- 10% of the ocean now protected. We have four years to triple it (2026) – Oceanographic
- Dominica creates the world’s first Sperm Whale Reserve, a win for conservation and climate (2025) – One Earth
- Dominica Establishes World’s First Sperm Whale Reserve, a Boost for Climate, Biodiversity and the Local Economy (2023) – National Geographic Society
- Extensive Use of Habitat-Damaging Fishing Gears Inside Habitat-Protecting Marine Protected Areas (2022) – Frontiers in Marine Science
- The Important Marine Mammal Area Network: A Tool for Systematic Spatial Planning in Response to the Marine Mammal Habitat Conservation Crisis (2022) – Frontiers in Marine Science
- The MPA Guide: A framework to achieve global goals for the ocean (2021) – Science
- Elevated trawling inside protected areas undermines conservation outcomes in a global fishing hot spot (2018) – Science
- Place-based approaches to marine mammal conservation (2016) – Aquatic Conservation
- Dynamic ocean management: Defining and conceptualizing real-time management of the ocean (2015) – Marine Policy
- Global conservation outcomes depend on marine protected areas with five key features (2014) – Nature
- The Importance of Marine Protected Areas (MPAs) – National Geographic Education
- MPAtlas – Marine Conservation Institute
- The MPA Guide – MPAtlas
Lesson 4 · Responsible whale watching
Articles and studies
- Your guide to the best, most responsible whale watching destinations (2026) – Travel Counsellors
- What is responsible whale watching? – IWC Whale Watching Handbook
- Importance of whale watching: Protecting marine life – IFAW
- Responsible whale watching and whale welfare – ResearchGate
- Keeping whales in captivity is wrong – Responsible Travel
- Wild vs. Captivity – Animal Welfare Institute
Lesson 5 · Citizen science
Articles and studies
- From citizen science to AI models: Advancing cetacean vocalization automatic detection through multi-annotator campaigns (2024) – Ecological Informatics
- The WhaleReport Alert System: Mitigating threats to whales with citizen science (2024) – Biological Conservation
- Population changes in a whale breeding ground revealed by citizen science noninvasive genetics (2022) – Global Ecology and Conservation
- Cidadãos, mídia social e ciência: a Ciência Cidadã como ferramenta de engajamento público na pesquisa com cetáceos – Maricato et al. (in Portuguese)
- Happywhale – Happywhale
- OceanWatchers – ORCA
- Citizen science – Whale Wise
- Whale Report Alert System (WRAS) – Ocean Wise
The Deep Dive
The depth records on the Deep Dive page.
- Mechanics, hydrodynamics and energetics of blue whale lunge feeding: efficiency dependence on krill density (2011) – Goldbogen et al., Journal of Experimental Biology 214: 131–146 · blue whale, 52–315 m
- Freediving – no limit (men) – Guinness World Records · Herbert Nitsch, 214 m, 2007
- Respiration and deep diving in the bottlenose porpoise (1969) – Ridgway, Scronce & Kanwisher, Science 166: 1651–1654 · bottlenose dolphin
- How deep can dolphins dive? – Dolphin Communication Project · bottlenose dolphin
- Movement and diving of killer whales (Orcinus orca) at a Southern Ocean archipelago (2015) – Reisinger et al., Journal of Experimental Marine Biology and Ecology 473: 90–102 · orca
- Deep-diving foraging behaviour of sperm whales (Physeter macrocephalus) (2006) – Watwood et al., Journal of Animal Ecology 75: 814–825 · sperm whale
- How do deep-diving sea creatures withstand huge pressure changes? (2002) – Ponganis & Kooyman, Scientific American · sperm whale, ~2,000 m
- Using narwhals as oceanographic sampling platforms in the high Arctic (2007) – Laidre & Heide-Jørgensen, Oceanography 20(4) · narwhal
- Deep-diving by narwhals Monodon monoceros: differences in foraging behavior between wintering areas? (2003) – Laidre et al., Marine Ecology Progress Series 261: 269–281 · narwhal
- First long-term behavioral records from Cuvier’s beaked whales (Ziphius cavirostris) reveal record-breaking dives (2014) – Schorr et al., PLOS ONE 9(3): e92633 · 2,992 m
- Longest dive by a mammal – Guinness World Records · Cuvier’s beaked whale, 222 minutes (Quick et al. 2020, Journal of Experimental Biology)
Images & sounds
Photographers, recordists and licences.
Sound recordings
- NOAA. Cornell University, Cornell Lab of Ornithology, K. Lisa Yang Center for Conservation Bioacoustics. 2023. Blue Whale. NOAA Sounds of the Ocean – used in “The blue whale”
- NOAA. NOAA Fisheries, NEFSC, Passive Acoustics Branch. 2023. Humpback Whale. NOAA Sounds of the Ocean – used in “The humpback whale”
- NOAA. NOAA Fisheries, NEFSC, Passive Acoustics Branch. 2023. North Atlantic Right Whale 1. NOAA Sounds of the Ocean – used in “Right whales”
- NOAA. NOAA Fisheries, NEFSC, Passive Acoustics Branch. 2023. Bottlenose Dolphin. NOAA Sounds of the Ocean – used in “Dolphins”
- NOAA. NOAA Fisheries, NEFSC, Passive Acoustics Branch. 2023. Sperm Whale. NOAA Sounds of the Ocean – used in “Sperm whale”
- NOAA. NOAA Fisheries, NEFSC, Passive Acoustics Branch. 2023. Cuvier’s Beaked Whale. NOAA Sounds of the Ocean – used in “Beaked whales”
- NOAA. NOAA Fisheries, NEFSC, Passive Acoustics Branch. 2023. Pilot Whale. NOAA Sounds of the Ocean – used in “Pilot whales”
- NOAA. Scottish Association for Marine Science, Denise Risch. 2023. Harbor Porpoise. NOAA Sounds of the Ocean – used in “Porpoises”
- NOAA. Alfred Wegener Institute, Ilse Van Opzeeland. 2023. Killer Whale. NOAA Sounds of the Ocean – used in “Orca”
Photographs
- Photo: Andriy Nekrasov – Plastic and chemical pollution
- Photo: Ethan Daniels – Pilot whales
- Photo: Francesco Ungaro – Underwater living
- Photo: Hernan Segui – Migration
- Photo: Ivan Stecko – Underwater living
- Photo: NOAA Fisheries – Porpoises
- Photo: NOAA Fisheries, Southwest Fisheries Science Center – Whaling
- Photo: NOAA Photo Library, CC BY 2.0 – Beaked whales
- Photo: Paula Olson, NOAA – The vaquita
- Photo: Roland Edler, CC BY 4.0 – Beaked whales
- Photo: Vicki Beaver, NOAA Fisheries – The bowhead whale
- Photo: marnock, Pexels – Porpoises
- Adobe Stock – Mammals that chose the ocean
- Shutterstock – 28 lessons
- Photo: Adam Ernster, Pexels – orca breaching, home page
Diagrams
- Diagrams and charts in the lessons were made for this course.